Here is a post copied from a friend on Trip Advisor. She has never toured with any of these guides but has seem them all along the Northern Range road/Lamar Valley when she is in the park. She lives in Billings so goes to the park at least monthly during May-October.
"There are lots of great private wildlife tour companies that operate out of Gardiner. I don't have a reason to take a private wildlife tour, but have had the opportunity to observe many of them in action during my multiple trips to Yellowstone Park each year. The following are ones I would recommend that you look at to see if they fit your interests and needs.
Yellowstone Guide (Jort Vanderveen)
https://yellowstoneguide.org/
https://www.facebook.com/yellowstoneguide/
Yellowstone Wild (Emil McCain)
https://www.yellowstonewildtours.com/tours/
https://www.facebook.com/YellowstoneWild/
Yellowstone Wolf Tracker (Nathan Varley and Linda Thurston)
https://www.wolftracker.com/index.php
https://www.facebook.com/yellowstonewolftracker/
Yellowstone Insight (MacNeil Lyons)
http://yellowstoneinsight.com/
https://www.facebook.com/YellowstoneInsight/"
Yellowstone Forever is the non-profit organization that supports the educational mission of YNP. I've done a number of their Field Seminars over the years and they are quality programs. They were specialist programs focused on either history or birding so I've never done one of their general programs. They do have some wildlife programs a couple of days a week during the summer season.
https://www.yellowstone.org/summer-yellowstone-day-adventures/
There are also Wake up to Wildlife tours out of Mammoth to Lamar Valley run by Xanterra the lodging provider. These would be fine if they are more in your budget and they do run every day.
https://www.yellowstonenationalparklodges.com/adventure/land-adventures/wake-up-to-wildlife/
The wildlife guides are all on radios so they hear what others are spotting and where. The tours don't go any place you can't go on your own. They stay on the paved road or there is an occasional foray down a couple of dirt/gravel roads.
TIMING: I would recommend you do a wildlife tour your first morning after arriving the day before. IF you are not used to spotting wildlife this will help you know where and how to look. It will be early so it depends on where you are traveling from as to whether this will work for you. I'd definitely do it out of Mammoth as this will be to the wildlife-rich area of Lamar Valley. The next day when you are transiting to Old Faithful you can re-drive the same area you covered and see what you can see. Backtracking is not a problem in Yellowstone. When I am staying on the North end, I drive the Northern Range (Mammoth to the NE entrance Road and out the NE entrance road almost to the NE entrance) every day or sometimes twice a day.
RANGER PROGRAMS:
Here also is a link to the 2025 Ranger programs. I have no idea if these will be the same or not for 2026 and they probably don't even know yet. Rangers start arriving to Old Faithful and Canyon for the opening day of the interior of the park which is the 3rd Friday of April and continue to arrive in batches up to the real beginning of the summer season on Memorial Day weekend. They then start leaving in batches, usually Aug 31, Sept 30 and the last ones turn out the lights on October 31.
https://www.nps.gov/yell/upload/2025-YELL-Summer-Ranger-Program-Schedule-Tabloid-11-x-17.pdf
This page should give you a link to the 2026 programs when they are uploaded sometime in April or by mid-May.
https://www.nps.gov/yell/planyourvisit/ranger-programs.htm